Species code:	CHLPD
Taxonomy:	Bacteria; Chlorobi; Chlorobia; Chlorobiales; Chlorobiaceae; Chlorobium/Pelodictyon group; Chlorobium (TaxID: 290317) [NEWT/ NCBI]
Description:	Chlorobium phaeobacteroides strain DSMZ 266 T is a non-motile, rod-shaped green sulphur bacterium (GSB) that can form elongated cells. GSB are commonly found in illuminated, stratified, and anoxic aquatic environments, sediments, and other sulfide-rich environments including hot springs. Because of unique adaptations of their light-harvesting antennae, these bacteria are capable of growth at light intensities under which no other phototrophs can survive. GSB are strictly anaerobic and obligately photoautotrophic in growth mode, and none is capable of dark respiratory or strictly fermentative metabolism. Most of these bacteria use electrons derived from reduced sulfur compounds in combination with light energy to reduce carbon and nitrogen. Many GSB strains can also utilize hydrogen as an electron donor for photoautotrophic growth. Unlike in most photosynthetic organisms, carbon fixation in the GSB occurs by the reverse TCA cycle reactions. The photosynthetic apparatus includes homodimeric type I reaction centers that are distantly related to photosystem I reaction centers of higher plants and cyanobacteria. The light harvesting antenna, chlorosomes, are ovoid bodies surrounded by a lipid monolayer and filled with more than 200,000 BChl e molecules and ~2500 BChl a molecules associated with the protein CsmA . This type strain was isolated from the anoxic sulfide containing water 19.5 m below surface of meromictic Lake Blankvann in Norway and is a representative of the brown-colored GSB species. The cells contain BChl e and BChl a as well as isorenieratene and b -isorenieratene as the major photosynthetic pigments. The strain requires vitamin B 12 for growth and lacks assimilatory sulfate reduction. Acetate and fructose can be assimilated during mixotrophic growth. Although it tested positive for hydrogenase activity, the strain cannot be grown with hydrogen as electron-donating substrate. Chl. phaeobacteroides lacks gas vesicles and grows in freshwater medium (adapted from http://genome.jgi-psf.org/finished_microbes/chlph/chlph.home.html).
Properties:	Presence of flagella: No Interaction: No Number of membranes: 2 Number of inteins:0
Statistics:	Number of CHLPD entries in the UniProt Knowledgebase: 2579 (237 in UniProtKB/Swiss-Prot + 2342 in UniProtKB/TrEMBL)

Genome structure:	Chromosome EMBL accession number CP000492 (circular; 3,133,902 bp) (download entry) (download Genome Reviews).
Reference(s):	[1] Copeland A., Lucas S., Lapidus A., Barry K., Detter J.C., Glavina del Rio T., Hammon N., Israni S., Pitluck S., Goltsman E., Schmutz J., Larimer F., Land M., Hauser L., Mikhailova N., Li T., Overmann J., Bryant D.A., Richardson P. ; "Complete sequence of Chlorobium phaeobacteroides DSM 266."; Submitted (DEC-2006) to the EMBL/GenBank/DDBJ databases.
Web links:	Official genome site(s): http://genome.jgi-psf.org/finished_microbes/chlph/chlph.home.html Other web sites: Entrez Genome Project: http://www.ncbi.nlm.nih.gov/sites/entrez?Db=genomeprj&Cmd=DetailsSearch&Term=txid290317%5Borgn%5D GIB (DDBJ): http://gib.genes.nig.ac.jp/single/index.php?spid=Cpha_DSM266 PEDANT: http://pedant.gsf.de:3345/pedant3htmlview/pedant3view?Method=start_method&Db=p3_p12609_Chl_phaeo  EBI Proteome Analysis page

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